 
          3072
        
        
          Proceedings of the 18
        
        
          th
        
        
          International Conference on Soil Mechanics and Geotechnical Engineering, Paris 2013
        
        
          
            Proceedings of the 18
          
        
        
          
            th
          
        
        
          
            International Conference on Soil Mechanics and Geotechnical Engineering, Paris 2013
          
        
        
          Figure 5. Relationships between suction and individual densities of the
        
        
          sand bubble crab and the decapod crustacean at three intertidal flats
        
        
          consistent with the ecohabitat chart showing that the burrowing
        
        
          capability of
        
        
          
            C. erythraeensis
          
        
        
          was markedly higher than other
        
        
          species and thereby exhibited the characteristics as the initial
        
        
          species inhabiting the reclaimed soils.
        
        
          Figure 7 shows the relationship between suction and the
        
        
          number of species that was obtained three years after the
        
        
          reclamation at the Tokuyama artificial intertidal flat. For the
        
        
          purpose of comparison, the results of the field surveys
        
        
          conducted fifteen years after the reclamation at another artificial
        
        
          intertidal flat, the Onomichi flat, are also plotted in this figure.
        
        
          One can observe that there is a close correlation between the
        
        
          diversity of species and suction. This fact is in agreement with
        
        
          the ecohabitat chart demonstrating that the suction-induced
        
        
          geoenvironment governs the manifestation of suitable and
        
        
          critical conditions for the diverse species, and the number of
        
        
          species, which can adapt to a severer geoenvironment, decreases
        
        
          with increasing suction and shear strength. As such, these
        
        
          results indicate that the difference in the suitable and critical
        
        
          geoenvironment among species contributes significantly to the
        
        
          distributions of the diverse species inhabiting there.
        
        
          4      CONCLUSION
        
        
          In the present study, we investigated systematically the linkage
        
        
          between the waterfront geoenvironment and the burrowing
        
        
          activity of six species of invertebrates in intertidal flats through
        
        
          a series of controlled laboratory experiments on the benthos-soil
        
        
          systems. The experimental results elucidate for the first time
        
        
          that there exist both suitable and critical geoenvironmental
        
        
          conditions for the burrowing activities of the diverse species
        
        
          irrespective of burrowing types, growth stages and weights. On
        
        
          the basis of these results, we have developed an ecohabitat chart
        
        
          which reveals complex interrelationships between such suitable
        
        
          and critical geoenvironment among species.
        
        
          In order to clarify the validity ofthe chart, weperformed
        
        
          integrated field observations, surveys and analyses concerning
        
        
          the waterfront geoenvironment and the species distributions at
        
        
          five natural and artificial intertidal flats. The results demonstrate
        
        
          that the way and where the diverse species lived are well
        
        
          consistent with the ecohabitat chart developed in this study.
        
        
          Overall, these results succeed not only in answering the
        
        
          fundamental question of why intertidal flats foster a complex
        
        
          ecosystem by the diverse species, from a view point of
        
        
          Ecological Geotechnics, but also establishing a new rational
        
        
          basis which can facilitate the conservation and restoration of
        
        
          habitats with rich natural ecosystems in intertidal zones.
        
        
          5     REFERENCES
        
        
          COP10. 2010. Strategic Plan for Biodiversity 2011-2020, tenth meeting
        
        
          of the Conference of the Parties (COP10).
        
        
          Kuwae T., Miyoshi E., Sassa S. and Watabe Y. 2010. Foraging mode
        
        
          shift in varying environmental conditions by dunlin
        
        
          
            Calidris alpine
          
        
        
          .
        
        
          
            Marine Ecology Progress Series
          
        
        
          406, 281-289.
        
        
          Sassa S. and Watabe Y. 2007. Role of suction dynamics in evolution of
        
        
          intertidal sandy flats: Field evidence, experiments, and theoretical
        
        
          model.
        
        
          
            Journal of Geophysical Research
          
        
        
          112, F01003,
        
        
          doi:10.1029/2006JF000575.
        
        
          Sassa S. and Watabe Y. 2008. Threshold, optimum and critical
        
        
          geoenvironmental conditions for burrowing activity of sand bubbler
        
        
          crab,
        
        
          
            Scopimera globosa. Marine Ecology Progress Series
          
        
        
          354, 191-
        
        
          199.
        
        
          Sassa S. and Watabe Y. 2009. Ecological Geotechnics: Performance of
        
        
          benthos activities controlled by suction, voids and shear strength in
        
        
          tidal flat soils.
        
        
          
            Proc. 17th Int. Conf. on Soil Mechanics and
          
        
        
          
            Geotechnical Engineering
          
        
        
          , ISSMGE, Egypt, 316-319.
        
        
          Sassa S., Watabe Y., Yang S. and Kuwae T. 2011. Burrowing Criteria
        
        
          and Burrowing Mode Adjustment in Bivalves to Varying
        
        
          Geoenvironmental Conditions in Intertidal Flats and Beaches.
        
        
          
            PLoS ONE
          
        
        
          6, No.9: e25041, doi:10.1371/journal.pone.0025041.
        
        
          (a)
        
        
          (b)
        
        
          Figure 6. Relationships between suction and densities of Arthropoda,
        
        
          Mollusca, and Annelida at the Tokuyama artificial intertidal flat
        
        
          Figure 7. Relationships between suction and diversity of species at
        
        
          two different artificial intertidal flats